Yes, 390mm is the number i'm getting from the axle center to th bottom of the arch brace (at centerline of fork). I took the wheel off to be sure.My Bluto is roughly 390mm from the axle center to the lowest point on the fork arch.
The radius of the XR4 is roughly 387 mm, inflated to 20 psi - will likely ride them around 12-15.
Upon closer inspection, the fit is a little tighter than I originally thought when looking at it while standing. When I actually look at the clearance from the lowest point on the center of the arch to the tread, it’s only about several mm.
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I knew many of the 29 x 3.0 are big including the XR4. I hear it is one of the larger ones. I didn't want to drop two hundo on tires and have a clearance issue.
I have only seen pics of other tires. Surprised TREK doesn't have a tire geometry page. Ridiculous.
Thanks again, Mark!